[Nucleic Acids]

Chapter 16 :The Lac Operon

A Model of How a Gene May be Regulated


The DNA within each cell contains thousands of genes. However, at any given time, only a small percentage of the genes are having their code being transcribed. Generally, only those genes which have to do with the particular cell type are transcribed on a regular basis within that cell.

As an example, for skin cells, only the genes having to do with making the cell a skin cell are transcribed. Those genes having to do with making nerve or blood cell proteins are "switched" off!

How then do the genes on a strand of DNA know when to be on or off?


Repressors: 'OFF' switches

Only genes that are directly needed are transcribed, others are ignored.

Lac Operon: Section of DNA with a promoter/operator in bacteria (one kind of prokaryitic cell) that acts as an on/off switch for several genes that control the breakdown of the sugar lactose into glucose + galactose.

Promoter: DNA section where RNA polymerase binds.

Operator: DNA section where repressor binds

Structural gene: Gene that makes an enzyme or other protein

Regulator gene: A gene not found next to the operon that makes a repressor protein.

Repressor: protein that shuts off operon. If bound to operator, it prevents RNA polymerase from initiating transcription.

Negative feedback system

*Repressor protein (always present) binds to operator.

*RNA polymerase is not allowed to move to structural gene as the repressor blocks its path.

*mRNA transcription is stopped, and thus enzyme production stops.

*This feedback system occurs when lactose concentration in cell is low.


Activators: 'ON' switches

Transcriptional activator: binds to DNA, holds RNA polymerase against promoter so transcription occurs.

*Activator protein binds to promoter. The activator is partly a molecule found only when energy is needed (because glucose levels are low).

*RNA polymerase now stays on promoter.

*RNA polymerase transcribes the structural genes

Positive Feedback & the Lac operon

*Activator (lactose) binds to repressor, changes its shape & it pops off operator.

*RNA polymerase binds to promoter and can now move past operator.

*RNA polymerase transcribes mRNA from structural gene.

*mRNA moves to ribosome where translation synthesizes the enzyme.

*The enzyme now breaks down the abundant lactose to glucose + galactose.



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