[Holboell's Rockcress] - [Rockcresses: The Genera Arabis, Boechera and Turritis West of the Cascade Mountains of Oregon and Washington]

Holboell's Rockcress, Second Rockcress, Secund Rockcress

Arabis holboellii var. retrofracta

Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis

Calyx hairs of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

The photo above shows a close-up view of the flowers of Holboell's rockcress (variety retrofracta) as seen along FS Rd 4650 to the east of Table Rock in the northern Wallowa-Whitman NF.........June 26, 2008. Note the numerous starlike hairs (many held aloft by short hairs) on the calyx lobes.

Revolute leaf edges of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)The photo at right shows the leaves and stems of Holboell's rockcress (variety retrofracta) as seen along FS Rd 21 above Murderers Creek, Malheur NF........5/24/2003.

Characteristics:

Holboell's rockcress is a biennial or short-lived perennial wildflower. One to several simple (to branched above) stems arise to a height of 100 cm. The lower part of the plant is strongly haired with simple or radially branched hairs, while the herbage is smooth to uniformly hairy above. The basal leaves are typically arranged in a basal rosette of linear-oblanceolate to broad, oblanceolate-spatulate leaves which are 1-5.5 cm long and 2-7 mm wide. The blades range from entire to toothed and are typically covered with fine felt-like hairs or longer, coarse or fine hairs. The numerous stem leaves often overlap each other and are often petiolate, although the upper leaves on some varieties are clasping with heart-shaped bases. They range from lanceolate to narrowly oblanceolate in shape and 1-5 cm long and 2-7 mm wide. Their blades are typically pubescent although they may be glabrous on some varieties.

The inflorescence is a densely-flowered raceme of 20-100 white, pink or most commonly, pinkish-purple flowers. The pedicels are 5-15 mm long and either uniformly curved downwards or more commonly abruptly bent downwards at the base. The pedicels vary from glabrous to hairy. Each of the 4 sepals is glabrous to hairy and each is 2.5-5 mm long. Each of the 4 petals is 5-10 mm long. The fruits are glabrous siliques from 3-7 cm long and 1-2 mm wide.

Variety retrofracta has simple to branched stems above measuring 20-90 cm high. They have woolly-haired stems below and range from pubescent to nearly glabrous above. The basal leaves are greater than 3 mm wide with entire to toothed margins. The leaf margins of the stem leaves are noticeably rolled under. The pedicels are sharply bent downwards at their bases.


Varieties:

Variety collinsii: Stem leaves with heart-shaped bases that often clasp the stem. Petals less than 7 mm long. Pedicels bent sharply downwards at their base. The siliques are more or less straight. Found from Washington to Alberta, Wyoming and east through he Dakotas and Manitoba to Quebec.

Variety pendulocarpa: Stem leaves lack heart-shaped bases. Found from British Columbia south to California and east to Montana and Colorado.

Variety pinetorum: Stem leaves with heart-shaped bases that often clasp the stem. Basal leaves are coarsely hairy. Pedicels uniformly curved outwards and downwards, not sharply reflexed downwards at the base. The siliques are thus spreading-drooping with the tip often incurved. Found from British Columbia south to California and east to Saskatchewan and Colorado.

Variety retrofracta: Stem leaves with heart-shaped bases that often clasp the stem. Leaves with margins rolled under, the upper ones pubescent. The basal leaves are covered in short, dense, felt-like hairs. Pedicels bent sharply downwards at their base. Petals over 7 mm long. Siliques 1.5-2 mm wide and more or less straight. Found from the Yukon and British Columbia south to California and east to Alberta (and occasionally to Quebec) and Colorado.


Habitat:

Hairy rock cress is found in forests and forest openings at all elevations.


Range:

Hairy rock cress is found from the Arctic south through much of North America to California, Arizona, and New Mexico, and as far east as the mid-Atlantic states.


Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

What appears to be Boechera retrofracta in bloom along Forest Service Road 35 on Table Mountain, Wenatchee National Forest........June 4, 2013.

Upper stem leaves and inflorescence of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Lower stem leaves of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

The photo above shows a close-up view of the inflorescence and upper stem leaves of Holboell's rockcress (variety retrofracta) as seen along FS Rd 4650 to the east of Table Rock in the northern Wallowa-Whitman NF.........June 26, 2008.

Inflorescence of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Inflorescence of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

Close-ups of the inflorescence of Holboell's rock cress (var. retrofracta). Note the glabrous, spreading to reflexed pedicels and the numerous star-shaped hairs on the calyces. Photographed on Wedge Mt., Wenatchee National Forest.......June 8, 2009.

Stem leaves of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Inflorescence of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis) - Inflorescence of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

Additional close-up views of Holboell's rockcress as seen at the Lawrence Memorial Grasslands several miles south of Shaniko, OR.............May 15, 2010.

Pendulous seed pods of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

The photo above shows Holboell's rockcress (variety retrofracta) as seen along FS Rd 21 above Murderers Creek, Malheur NF........5/24/2003.

Pendulous seed pod of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

The photo above shows the fruit of Holboell's rockcress (variety retrofracta) as seen along FS Rd 21 above Murderers Creek, Malheur NF........5/24/2003. Note the pedicel which is curved sharply downwards.

Basal leaves of Holboell's Rockcress, Second Rockcress, Secund Rockcress: Arabis holboellii var. retrofracta (Synonyms: Arabis arcuata, Arabis caduca, Arabis consanguinea, Arabis dacotica, Arabis exilis, Arabis holboellii, Arabis holboellii var. collinsii, Arabis holbellii var. consanguinea, Arabis holboellii var. secunda, Arabis holbellii var. tenuis, Arabis kochii, Arabis lignipes, Arabis mcdougalii, Arabis polyantha, Arabis retrofracta, Arabis retrofracta var. multicaulis, Arabis rhodanthus, Arabis secunda, Arabis tenuis, Boechera exilis, Boechera holbellii var. retrofracta, Bochera hoboellii var. secunda, Boechera retrofracta, Boechera tenuis)

The photo above shows the basal leaves and lower stem leaves of Holboell's rockcress (variety retrofracta) as seen along FS Rd 21 above Murderers Creek, Malheur NF........5/24/2003.

Paul Slichter