Leathery Grapefern
Botrichium multifidum

The photo above shows a close-up of the broad leaf of leathery grapefern as seen along the trail to Willard Springs in Conboy Lake National Wildlife Refuge...................April 13, 2007.

The photo above shows the sporophore or fertile frond of leathery grapefern which consists of a panicle of sporangia which will release spores when mature. Photographed at Willard Springs in Conboy Lake NWR...................August 17, 2007.

The photo above shows the sporophore or fertile frond of leathery grapefern as well as the sterile frond laying upon the ground. Photographed at Willard Springs in Conboy Lake NWR...................August 17, 2007.
Paul Slichter