[Grapeferns and Moonworts of Mt. Adams Country]

Leather Grapefern, Leather Grape-fern, Leathery Grapefern

Sceptridium multifidum

Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida

Sterile frond of Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida)

The photo above shows a close-up of the broad leaf of leathery grapefern as seen along the trail to Willard Springs in Conboy Lake National Wildlife Refuge..........April 13, 2007.

Fertile frond of Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida)

The photo above shows the sporophore or fertile frond of leathery grapefern which consists of a panicle of sporangia which will release spores when mature. Photographed at Willard Springs in Conboy Lake NWR.........August 17, 2007.

Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida) - Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida)

Leathery grapeferns were numerous, here seen growing with broadpetal strawberry (Fragaria virginiana ssp. glauca) in the drier grassy meadows around the northern edge of Muddy Meadows, Mt. Adams Wilderness......July 13, 2018.

Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida)

The photo above shows the sporophore or fertile frond of leathery grapefern as well as the sterile frond laying upon the ground. Photographed at Willard Springs in Conboy Lake NWR.........August 17, 2007.

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Several photos of leather grapefern as seen on grassy, dry soils at a slightly higher elevation than the vernally moist meadows at the trailhead for the Muddy Meadows Trail #13 on the northern slopes of Mt. Adams. The fertile part of the leaf blade is shown with the evident, grape-like sporangia which gives this genus its name of "grapefern". No sterile blade was noticed. The fertile blade itself is about 10 cm long.

Leather Grapefern, Leather Grape-fern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium,  Botrychium silaifolium, Osmunda multifida)

Leathery grapefern as seen along the Willard Springs Trail, Conboy Lake National Wildlife Refuge........September 17, 2017.

Paul Slichter