[Swordferns: The Genus Polystichum East of the Crest of the Cascade Mountains of Oregon and Washington] - [Ferns East of the Crest of the Cascade Mountains of Oregon and Washington]

Holly fern as seen along the Canyon Mountain Trail #218, Strawberry Mountain Wilderness........August 19, 2011.

Close-ups of the dorsal and ventral surfaces of the leaflets of holly fern as seen along the Canyon Mountain Trail #218, Strawberry Mountain Wilderness.........August 19, 2011.

Several views of holly fern as seen on cliffs about one-half mile north of Wildhorse Lake, Steens Mountain, Harney County, Oregon............July 10, 2014.

Holly fern along the East Fork Lostine River Trail on a series of switchbacks with forest to the south and a granitic boulder field to the north in the Eagle Cap Wilderness.........August 4, 2016.