[Grapeferns and Moonworts East of the Crest of the Cascade Mountains of Oregon and Washington] - [Ferns East of the Cascade Crest of Oregon and Washington]

Leather Grapefern, Leathery Grapefern

Sceptridium multifidum

Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium, Botrychium silaifolium, Osmunda multifida

Sterile frond of Leather Grapefern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium Botrychium silaifolium, Osmunda multifida)

The photo above shows a close-up of the broad leaf of leathery grapefern as seen along the trail to Willard Springs in Conboy Lake National Wildlife Refuge.........April 13, 2007.

Fertile frond of Leather Grapefern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium Botrychium silaifolium, Osmunda multifida)

The photo above shows the sporophore or fertile frond of leathery grapefern which consists of a panicle of sporangia which will release spores when mature. Photographed at Willard Springs in Conboy Lake NWR........August 17, 2007.

Leather Grapefern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium, Botrychium silaifolium, Osmunda multifida) - Leather Grapefern, Leathery Grapefern: Sceptridium multifidum (Synonyms: Botrychium californicum, Botrychium coulteri, Botrychium multifidum, Botrychium multifidum ssp. silaifolium, Botrychium silaifolium, Osmunda multifida)

Leathery grapefern as photographed at the dry upper edge of moist meadows at an old reservoir about one mile above Olive Lake, Umatilla National Forest..........July 28, 2013.
Paul Slichter